'Extraordinary' sign language gorilla Koko dies in California

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An "extraordinary" gorilla that mastered sign language has died.
The Gorilla Foundation announced on Twitter that it was "sad to announce the passing of our beloved Koko".
The foundation said the western lowland gorilla, who developed an "extraordinary mastery" of signing, died in her sleep aged 46.
Koko featured in documentaries and appeared on the cover of National Geographic twice after starting to learn sign language when she was a year old.
Animal psychologist Francine Patterson and Koko in 1972
She was born at San Francisco Zoo but three years later was moved to Stanford, where under the tutorship of Dr Francine Patterson and Dr Ronald Cohn she developed her skills.
Koko was later joined in the project by a second lowland gorilla, Michael, and was eventually moved to the Santa Cruz mountains in California where they were joined by a third gorilla, Ndume.

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